The Let’s Stay Together celebrity campaign, backed by Simon Cowell and Davina McCall spent £133,832.
Other No backers included The No Borders Campaign which shelled out £147,510 and the Working for Scotland group shelled out £118,303, while the Orange Lodge was another big donor, spending £47,072 in the last 16 weeks of the campaign.
On the Yes side, Business for Scotland spent £143,027 trying to woo industry towards independence, while the left wing 1001 campaign spent £72,055 and comedy club impresario Tommy Sheppard, now an SNP candidate in Edinburgh, for the general election, shelled out £35,094.
A number of groups including Labour for Independence and the Scottish Jacobite Party are now facing “enforcement action” after failing to submit returns, the Electoral Commission said.
In total, campaigners for independence spent £399,405, while campaigners for a No outcome reported spending of £1,034,933
Groups which spent between £10,000 and £250,000 were required to submit spending returns to the Commission by 18 December 2015.
Reported spending by campaigners
Groups backing Yes
1001 Campaign: £72,055
Business for Scotland: £143,027
Christians for Independence: £29,248
Generation Yes: £14,065
Mr Tommy Sheppard: £35,094
National Collective: £54,849
Scottish Green Party: £13,734
Scottish Socialist Party: £12,728
Women for Independence: £24,605
Groups backing No
Better With Scotland: £47,042
Communication Workers Union (CWU): £20,437
Cumbria Broadband Rural and Community Projects Limited: £65,921
Grand Orange Lodge of Scotland: £47,072
Let’s Stay Together: £133,832
Liberal Democrats: £187,585
Mr Angus MacDonald: £110,644
No Borders Campaign: £147,510
The Scottish Research Society: £82,202
Union of Shop, Distributive & Allied Workers (USDAW): £30,550
WFS2014 (Working for Scotland): £118,303
Total ‘Yes’: £399,405
Total ‘No’: £1,034,933
Total overall: £1,434,338