Steve Thompson drops to the bench where he is joined by Wasps lock Simon Shaw, who has replaced Dave Attwood after recovering from a calf injury.
Hartley made a dynamic impact off the bench against the All Blacks last weekend, scoring his first Test try to spark England's second-half comeback. The fiery 24-year-old was arguably fortunate not to have been cited for a challenge on Richie McCaw. But Hartley added to an impressive scrummaging performance and England will again be targeting the Wallabies' fragile set-piece.
England dominated Australia's scrum during the drawn summer Test series and they look a more formidable unit with Andrew Sheridan back in the side at loose-head prop. England manager Martin Johnson has decided against making changes to his centre combination, with Mike Tindall and Shontayne Hape continuing their partnership.
England team: B Foden (Northampton); C Ashton (Northampton), M Tindall (Gloucester), S Hape (Bath), M Cueto (Sale); T Flood (Leicester), B Youngs (Leicester); A Sheridan (Sale), D Hartley (Northampton), D Cole (Leicester), C Lawes (Northampton), T Palmer (Stade Francais), T Croft (Leicester), L Moody (Bath, capt), N Easter (Harlequins). Replacements: S Thompson (Leeds), D Wilson (Bath), S Shaw (Wasps), H Fourie (Leeds), D Care (Harlequins), C Hodgson (Sale), D Armitage (London Irish).